ORIGINAL ARTICLE
Lichenicolous fungi of the steppe zone of Ukraine: taxonomic diversity and phylogenetic notes
1
W. Szafer Institute of Botany, Polish Academy of Sciences, Lubicz
46, 31-512 Kraków, Poland
2
Kherson State University, Shevchenka 14, IvanoFrankivsk, 76018,
Ukraine
3
Ukrainian Botanical Society, Kulchytska 17, 47501, Posukhiv, Ternopil
region, Ukraine
4
V.N. Karasin Kharkiv National University, Svobody sq., 4, Kharkiv,
61022, Ukraine
5
Homilshanski Lisy National Nature Park, Monastyrska 27, Koropove
63523, Kharkiv Region, Ukraine
6
M.G. Kholodny Institute of Botany, National Academy of Science of
Ukraine, Tereshenkivska 2, Kyiv, 01601, Ukraine
Submission date: 2026-01-16
Acceptance date: 2026-03-26
Online publication date: 2026-09-04
Publication date: 2026-09-04
Plant and Fungal Systematics 2026; 71(2): 273-318
KEYWORDS
ABSTRACT
Lichenicolous fungi represent an essential, but still insufficiently studied, component
of the mycobiota of Europe. The steppe zone of Ukraine, despite its high ecological
heterogeneity and conservation importance, has remained particularly underexplored. In
this study, we provide a comprehensive revision of lichenicolous fungi occurring in the
steppe zone of Ukraine based on extensive field collections from the last decade and
a critical evaluation of published data. A total of 168 species are confirmed for the region,
including ten species reported here for the first time from Ukraine: Arthonia urceolata,
Endococcus verrucosus, Gonatophragmium lichenicola, Lichenostigma chlaroterae, Llanorella
ramalinae, Nesolechia fusca, Spirographa skorinae, Tremella celata, T. parietinae,
and Zyzygomyces physconiae. Molecular data further clarify the phylogenetic position of
Phoma candelariellae, which forms an isolated lineage within Phaeosphaeriaceae. We
introduce the new genus Phomella and provide a new combination, Phomella candelariellae.
Several previously unknown host associations are documented, including Calogaya
saxicola as a new host for Chaetopyrena penicillata; Cetraria aculeata for Cladosporium
licheniphilum; Circinaria hispida, Xanthoparmelia camtschadalis, and X. pokornyi for
Didymocyrtis cladoniicola; Xanthoria monofoliosa for Didymocyrtis epiphyscia; Xanthoparmelia
camtschadalis for both Epithamnolia xanthoriae and Nesolechia fusca; and
Xanthoria parietina for Protocreopsis globulosa. Plowrightia mereschkowskyi is considered
a new synonym for Arthonia urceolata.
FUNDING
This study was financially supported
by the Ministry of Science and Education of Ukraine (projects
0116U004735 and 0219U000270).
REFERENCES (149)
1.
Altschul, S. F., Gish, W., Miller, W., Myers, E. W. & Lipman, D. J. 1990. Basic local alignment search tool. Journal of Molecular Biology 215: 403–410. https://doi.org/10.1016/S0022-....
3.
Berger, F. & Brackel, W. von 2011. Eine weitere art von Phaeosporobolus auf Lecanora chlarotera. Herzogia 24: 351–356. https://doi.org/10.13158/heia.....
4.
Berger, F., Braun, U. & Heuchert, B. 2015. Gonatophragmium lichenophilum sp. nov. – a new lichenicolous hyphomycete from Austria. Mycobiota 5: 7–13. https://doi.org/10.12664/mycob....
5.
Bielczyk, U. R., Bylińska, E., Czarnota, P., Czyżewska, K., Guzow-Krzemińska, B., Hachułka, M., Kiszka, J., Kowalewska, A., Krzewicka, B., Kukwa, M., Leśniański, G., Śliwa, L. & Zalewska, A. 2005. Contribution to the knowledge of lichens and lichenicolous fungi of western Ukraine. Polish Botanical Journal 50: 39–64.
6.
Boberski, W. 1886. Systematischer Üebersicht der Flechten Galiziens. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien XXXVI: 243–286.
7.
Bodiak, H. 1907. Symbolae ad Mycologiam Haliciae orientalis. Fungi agri Berežanensis. Sammelschrift der Mathematisch-naturwissenschaftlich-ärztlichen Section der Ševčenko-Gesellschaft der Wissenschaften in Lemberg 11: 1–141.
8.
Boiko, T. O. 2008. The lichens and lichenicolous fungi of limestone outcrops of the Nature Reserve “Yelanetsky step”. Chornomorski Botanical Journal 4(1): 84–88.
9.
Bomar, M. T. & Knöpfel, S. A. 1992. A method for the estimation of the culturing quality of dehydrated mycological media. International Journal of Food Science and Technology 27(5): 589–592. https://doi.org/10.1111/j.1365....
10.
Brackel, W. von 2014. Kommentierter Katalog der flechtenbewohnenden Pilze Bayerns. Bibliotheca Lichenologica 109: 1–476.
11.
Braun, U., Khodosovtsev, A. Y., Darmostuk, V. V. & Diederich, P. 2016. Trichoconis hafellneri sp. nov. on Athallia pyracea and Xanthoria parietina, a generic discussion of Trichoconis and keys to the species of this genus. Herzogia 29: 307–314. https://doi.org/10.13158/heia.....
12.
Calatayud, V. & Etayo, J. 1999. Codonmyces and Lichenostella, two new genea of lichenicolous conidial fungi. The Lichenologist 36: 593–601.
13.
Calatayud, V. & Etayo, J. 2001. Five new species of lichenicolous conidial fungi from Spain. Canadian Journal of Botany 79: 223–230. https://doi.org/10.1139/b00-14....
14.
Calatayud, V., Navarro-Rosinés, P. & Hafellner, J. 2002. A synopsis of Lichenostigma subgen. Lichenogramma (Arthoniales), with a key to the species. Mycological Research 106: 1230–1242. https://doi.org/10.1017/S09537....
15.
Calatayud, V., Barreno, E. & Rico, V. J. 2004. Two lichenicolous species of Arthonia on vagrant Aspicilia species. Bibliotheca Lichenologica 88: 67–78.
16.
Calatayud, V., Navarro-Rosinés, P. & Hafellner, J. 2013. Contributions to a revision of Cercidospora (Dothideales), 2: Species on Lecanora s.l., Rhizoplaca and Squamarina. Mycosphere 4: 539–557. https://doi.org/10.5943/mycosp....
17.
Cezanne, R., Eichler, M., Berger, F., Brackel, W. von, Breuss, O., Dolnik, C., Dornes, A. P., Eckstein, J., Schultz, M., Thüs, H. & Türk, R. 2023. Neu- und Wiederfunde von Flechten, flechtenbewohnenden und flechtenähnlichen Pilzen in Deutschland. Herzogiella 10: 80–102.
18.
Chernov, V. K. 1895. About lichens of the city of Kharkov and its environs. Proceedings of the Society of Naturalists at Kharkov University 28: 209–249.
19.
Crous, P. W., Verkley, G. J. M., Groenewald, J. Z. & Samson, R. A. 2009. Fungal Biodiversity. [CBS Laboratory Manual Series no.1.]. CBS-KNAW Fungal Biodiversity Centre, Utrecht.
20.
Crous, P. W., Akulov, A., Balashov, S., Boers, J., Braun, U., Castillo, J., Delgado, M. A., Denman, S., Erhard, A., Gusella, G., Jurjević, Ž., Kruse, J., Malloch, D. W., Osieck, E. R., Polizzi, G., Schumacher, R. K., Slootweg, E., Starink-Willemse, M., Van Iperen, A. L., Verkley, G. J. M. & Groenewald, J. Z. 2023. New and Interesting Fungi. 6. Fungal Systematics and Evolution 11: 109–156. https://doi.org/10.3114/fuse.2....
21.
Crous, P. W., Wingfield, M. J., Jurjević, Ž., Balashov, S., Osieck, E. R., Marin-Felix, Y., Luangsa-ard, J. J., Mejía, L. C., Cappelli, A., Parra, L. A., Lucchini, G., Chen, J., Moreno, G., Faraoni, M., Zhao, R. L., Weholt, O., Borovička, J., Jansen, G. M., Shivas, R. G., Tan, Y. P., Akulov, A., Alfenas, A. C., Alfenas, R. F., Altés, A., Avchar, R., Barreto, R. W., Catcheside, D. E. A., Chi, T. Y., Esteve-Raventós, F., Fryar, S. C., Hanh, L. T. M., Larsbrink, J., Oberlies, N. H., Olsson, L., Pancorbo, F., Raja, H. A., Thanh, V. N., Thuy, N. T., Ajithkumar, K., Akram, W., Alvarado, P., Angeletti, B., Arumugam, E., Atashi Khalilabad, A., Bandini, D., Baroni, T. J., Barreto, G. G., Oertmann, D. B., Bose, T., Castañeda Ruiz, R. F., Couceiro, A., Cykowska-Marzencka, B., Dai, Y. C., Darmostuk, V., Da Silva, S. B. G., Dearnaley, J. D. W., De Azevedo Santiago, A. L. C. M., Declercq, B., De Freitas, L. W. S., De La Peña-Lastra, S., Delgado, G., De Lima, C. L. F., Dhotre, D., Dirks, A. C., Eisvand, P., Erhard, A., Ferro, L. O., García, D., García-Martín, A., Garrido-Benavent, I., Gené, J., Ghobad-Nejhad, M., Gore, G., Gunaseelan, S., Gusmão, L. F. P., Hammerbacher, A., Hernández-Perez, A. T., Hernández-Restrepo, M., Hofmann, T. A., Hubka, V., Jiya, N., Kaliyaperumal, M., Keerthana, K. S., Ketabchi, M., Kezo, K., Knoppersen, R., Kolarczyková, D., Kumar, T. K. A., Aesoe, T., Langer, E., Larsson, E., Lodge, D. J., Lynch, M. J., Maciá-Vicente, J. G., Mahadevakumar, S., Mateos, A., Mehrabi-Koushki, M., Miglio, B. V., Noor, A., Oliveira, J. A., Pereira, O. L., Piatek, M., Pinto, A., Ramírez, G. H., Raphael, B., Rawat, G., Renuka, M., Reschke, K., Ruiz Mateo, A., Saar, I., Saba, M., Safi, A., Sánchez, R. M., Sandoval-Denis, M., Savitha, A. S., Sharma, A., Shelke Sonawane, D., Souza, M. G. A. P., Stryjak-Bogacka, M., Thines, M., Thomas, A., Torres-Garcia, D., Traba, J. M., Vauras, J., Vermaas, M., Villarreal, M., Vu, D., Whiteside, E. J., Zafari, D., Starink-Willemse, M. & Groenewald, J. Z. 2024. Fungal Planet description sheets: 1697–1780. Fungal Systematics and Evolution 14: 325–577. https://doi.org/10.3114/fuse.2....
22.
Crous, P. W., Catcheside, D. E. A., Catcheside, P. S., Alfenas, A. C., Alfenas, R. F., Barreto, R. W., Lebel, T., Balashov, S., Broadbridge, J., Jurjević, Ž., Peña-Lastra, S. D. L., Hoffmann, R., Mateos, A., Riebesehl, J., Shivas, R. G., Soliz-Santander, F. F., Tan, Y. P., Altés, A., Bandini, D., Carriconde, F., Cazabonne, J., Czachura, P., Gryta, H., Eyssartier, G., Larsson, E., Pereira, O. L., Rigueiro-Rodríguez, A., Wingfield, M. J., Ahmad, W., Bibi, S., Denman, S., Esteve-Raventós, F., Hussain, S., Illescas, T., Luangsa-ard, J. J., Möller, L., Mombert, A., Noisripoom, W., Olariaga, I., Pancorbo, F., Paz, A., Piątek, M., Polman-Short, C., Suárez, E., Afshan, N. S., Ali, H., Arzanlou, M., Ayer, F., Barratt, J., Bellanger, J. ‐M., Bidaud, A., Bishop-Hurley, S. L., Bohm, M., Bose, T., Campo, E., Chau, N. B., Çolak, Ö. F., Cordeiro, T. R. L., Cruz, M. O., Custódio, F. A., Couceiro, A., Darmostuk, V., Dearnaley, J. D. W., De Azevedo Santiago, A. L. C. M., De Freitas, L. W. S., Yáñez-Morales, M. D. J., Domnauer, C., Dentinger, B., Dhileepan, K., De Souza, J. T., Dovana, F., Eberhardt, U., Eisvand, P., Erhard, A., Fachada, V., Garcia-Martín, A., Groenewald, M., Hammerbacher, A., Harms, K., Haroon, S., Haqnawaz, M., Henriques, S., Hernández, A. J., Jacobus, L. M., Jaen-Contreras, D., Jangsantear, P., Kaygusuz, O., Knoppersen, R., Kumar, T. K. A., Lynch, M. J., Mahiques, R., Maraia, G. L., Marbach, P. A. S., Mehrabi-Koushki, M., Miller, P. R., Mongkolsamrit, S., Moreau, P.‐A., Oberlies, N. H., Oliveira, J. A., Orlovich, D., Pérez-Méndez, A. S., Pinto, A., Raja, H. A., Ramírez, G. H., Raphael, B., Rodrigues, A., Rodrigues, H., Ramos, D. O., Safi, A., Sarwar, S., Saar, I., Sánchez, R. M., Santana, J. S., Scrace, J., Sales, L. S., Silva, L. N. P., Stryjak-Bogacka, M., Tacconi, A., Thanh, V. N., Thomas, A., Thuy, N. T., Toome, M., Valdez-Carrazco, J. M., Van Vuuren, N. I., Vasey, J., Vauras, J., Vila-Viçosa, C., Villarreal, M., Visagie, C. M., Vizzini, A., Whiteside, E. J. & Groenewald, J. Z. 2025. Fungal Planet description sheets: 1781–1866. Persoonia 54: 327–587. https://doi.org/10.3114/persoo....
23.
Darmostuk, V. V. 2016. The genus Cercidospora (Dothideales) in Ukraine. Ukrainian Botanical Journal 73: 262–267. https://doi.org/10.15407/ukrbo....
24.
Darmostuk, V. V. 2019a. Additions to the lichenicolous mycobiota of Ukraine: the genus Zwackhiomyces (Xanthopyreniaceae, Collemopsidiales). Ukrainian Botanical Journal 76: 301–315. https://doi.org/10.15407/ukrbo....
25.
Darmostuk, V. V. 2019b. The genus Lichenoconium (Lichenoconiaceae, Ascomycota) in Ukraine. Ukrainian Botanical Journal 76: 101–113. https://doi.org/10.15407/ukrbo....
26.
Darmostuk, V. V. 2021a. Pronectria gromakovae, a new lichenicolous fungus on Lecanora populicola and notes on other records from Kharkiv region (Ukraine). Lindbergia 44: linbg.01141. https://doi.org/10.25227/linbg....
27.
Darmostuk, V. V. 2021b. Lichenicolous fungi on Verrucaria s.lat. in Ukraine with the description of Zwackhiomyces khodosovtsevii sp. nov. and a key to the lichenicolous fungi on Verrucaria s.lat. Botanica Serbica 45: 293–301. https://doi.org/10.2298/BOTSER....
28.
Darmostuk, V. V. & Golovenko, Ye. O. 2016. Polycoccum aksoyi Halıcı & V. Atienza is a new species of lichenicolous fungi for Ukraine. Visnyk of Zaporizhzhya National University. Biological Sciences 1: 123–127.
29.
Darmostuk, V. V. & Khodosovtsev, A. Ye. 2017. Lichenicolous fungi of Ukraine: an annotated checklist. Studies in Fungi 2: 138–156. https://doi.org/10.5943/sif/2/....
30.
Darmostuk, V. V. & Khodosovtsev, A. Ye. 2019. Epibryon kondratyukii sp. nov., a new algicolous fungus, and notes on rare lichenicolous fungi collected in Southern Ukraine. Folia Cryptogamica Estonica 56: 109–116. https://doi.org/10.12697/fce.2....
31.
Darmostuk, V. V. & Khodosovtsev, A. Ye. 2020. Notes to lichen-forming and lichenicolous fungi in Ukraine I. Chornomorski Botanical Journal 16: 257–274. https://doi.org/10.32999/ksu19....
32.
Darmostuk, V. V. & Khodosovtsev, A. Ye. 2021. Old artificial parks as a key spot of corticolous lichen diversity in Southern Ukraine. Chornomorski Botanical Journal 17: 148–163. https://doi.org/10.32999/ksu19....
33.
Darmostuk, V. V. & Naumovich, G. O. 2016. Unguiculariopsis (Helotiaceae, Helotiales), a new genus for the mycobiota of Ukraine. Ukrainian Botanical Journal 73(4): 378–381. https://doi.org/10.15407/ukrbo....
34.
Darmostuk, V. & Sira, O. Ye. 2020. New and remarkable records of lichenicolous fungi from Ternopil Oblast (Ukraine). Czech Mycologia 72: 33–41. https://doi.org/10.33585/cmy.7....
35.
Darmostuk, V. & Sira, O. 2022. New and remarkable records of lichenicolous fungi from Ternopil Oblast (Ukraine). II. Folia Cryptogamica Estonica 59: 43–51. https://doi.org/10.12697/fce.2....
36.
Darmostuk, V. V., Khodosovtsev, A. Ye., Naumovich, G. O. & Kharechko, N. V. 2018. Roselliniella lecideae sp. nov. and other interesting lichenicolous fungi from the Northern Black Sea region (Ukraine). Turkish Journal of Botany 42: 354–361. https://doi.org/10.3906/bot-17....
37.
Darmostuk, V. V., Khodosovtsev, A. Ye., Gromakova, A. B., Sira, O. Ye., Davydov, D. A., Gavrylenko, L. M. & Khodosovtseva, Ya. A. 2021. Notes to lichen-forming and lichenicolous fungi in Ukraine II. Chornomorski Botanical Journal 17: 276–295. https://doi.org/10.32999/ksu19....
38.
Darmostuk, V. V., Khodosovtsev, A. Ye. & Kostikov, I. Yu. 2022. First lichenicolous records of Chaetopyrena penicillata. Mycotaxon 137: 591–602. https://doi.org/10.5248/137.59....
39.
Darmostuk, V. V., Khodosovtsev, A. Ye., Gromakova, A. B., Sira, O. Ye. & Bezsmertna, O. O. 2023. Notes to lichen-forming and lichenicolous fungi in Ukraine III. Chornomorski Botanical Journal 19: 58–75. https://doi.org/10.32999/ksu19....
40.
Darmostuk, V., Sira, O. & Govorun, O. 2024a. New data about overlooked terricolous lichen-forming and lichenicolous fungi of Ukraine. Folia Cryptogamica Estonica 61: 59–66. https://doi.org/10.12697/fce.2....
41.
Darmostuk, V., Gromakova, A. B., Kapets, N. V., Lavrynenko, K. V., Borovyk, D. V., Kuzemko, A. A. & Khodosovtsev, O. Ye. 2024b. Notes to lichen-forming and lichenicolous fungi in Ukraine IV. Chornomorski Botanical Journal 20: 19–35. https://doi.org/10.32999/ksu19....
42.
Darmostuk, V., Etayo, J., Rodriguez-Flakus, P., Kukwa, M. & Flakus, A. 2025. Expanding the genus Neobaryopsis (Calcarisporiaceae, Hypocreales): descriptions of two new species and two new combinations based on morphological and molecular data. MycoKeys 124: 291–308. https://doi.org/10.3897/mycoke....
43.
Didukh, Y. P. 2020. Biotopes of Steppe zone of Ukraine. DrukArt, Kyiv-Chernivtsi.
44.
Diederich, P. 2021. Notes on lichenicolous taxa of the asexual fungal genera Intralichen and Trimmatostroma, with a revised key and descriptions of four new species. Herzogia 34: 101–126.
45.
Diederich, P., Lawrey, J. D., Sikaroodi, M., van den Boom, P. & Ertz, D. 2012. Briancoppinsia, a new coelomycetous genus of Arthoniaceae (Arthoniales) for the lichenicolous Phoma cytospora, with a key to this and similar taxa. Fungal Diversity 52: 1–12. https://doi.org/10.1007/s13225....
46.
Diederich, P., Lawrey, J. D. & Ertz, D. 2018. The 2018 classification and checklist of lichenicolous fungi, with 2000 non-lichenized, obligately lichenicolous taxa. The Bryologist 121: 340–425. https://doi.org/10.1639/0007-2....
47.
Diederich, P., Millanes, A. M., Wedin, M. & Lawrey, J. D. 2022. Flora of lichenicolous fungi. Volume 1: Basidiomycota. National Museum of Natural History, Luxembourg.
48.
Diederich, P., Ertz, D. & Braun, U. 2024. Flora of Lichenicolous Fungi, Vol. 2, Hyphomycetes. National Museum of Natural History, Luxembourg.
49.
Diederich, P., Lawrey, J. D. & Ertz, D. 2025. The 2025 classification and checklist of lichenicolous fungi: documenting a rapidly growing knowledge of diversity. The Bryologist 128: 765–870. https://doi.org/10.1639/0007-2....
50.
Elenkin, A. 1901. Les lichens facultatifs. Bulletin du Jardin impérial botanique de St.-Pétersbourg 1: 129–154.
51.
Ertz, D., Diederich, P., Lawrey, J. D., Berger, F., Freebury, C. E., Coppins, B., Gardiennet, A. & Hafellner, J. 2015. Phylogenetic insights resolve Dacampiaceae (Pleosporales) as polyphyletic: Didymocyrtis (Pleosporales, Phaeosphaeriaceae) with Phoma-like anamorphs resurrected and segregated from Polycoccum (Trypetheliales, Polycoccaceae fam. nov.). Fungal Diversity 74: 53–89. https://doi.org/10.1007/s13225....
52.
Etayo, J. & Yazici, K. 2009. Microsphaeropsis caloplacae sp. nov. on Caloplaca persica in Turkey. Mycotaxon 107: 297–302. https://doi.org/10.5248/107.29....
53.
Fedorenko, N. M., Nadeina, O. V. & Kondratyuk, S. Ya. 2007. New and rare lichenicolous fungi from Ukraine. Ukrainian Botanical Journal 64(1): 47–56.
54.
Fernández-Brime, S., Gaya, E., Llimona, X., Wedin, M. & Navarro-Rosinés, P. 2020. Rhagadodidymellopsis endocarpi gen. et sp. nov. and Arthopyrenia symbiotica (Dothideomyceta), two lichenicolous fungi growing on Endocarpon species. Plant and Fungal Systematics 65: 176–184. https://doi.org/10.35535/pfsys....
55.
Flakus, A., Etayo, J., Miadlikowska, J., Lutzoni, F., Kukwa, M., Matura, N. & Rodriguez-Flakus, P. 2019. Biodiversity assessment of ascomycetes inhabiting Lobariella lichens in Andean cloud forests led to one new family, three new genera and 13 new species of lichenicolous fungi. Plant and Fungal Systematics 64: 283–344. https://doi.org/10.2478/pfs-20....
56.
Freire-Rallo, S., Diederich, P., Millanes, A. M. & Wedin, M. 2023. Five new species in the Tremella caloplacae complex. The Lichenologist 55: 223–239. https://doi.org/10.1017/S00242....
57.
Frisch, A., Holien, H., Klepsland, J. T., Suija, A. & Bendiksby, M. 2022. New data on lichenicolous fungi in Norway. Graphis Scripta 34: 59–133.
58.
Gardes, M. & Bruns, T. D. 1993. ITS primers with enhanced specificity for basidiomycetes – application to the identification of mycorrhizae and rusts. Molecular Ecology 2: 113–118. https://doi.org/10.1111/j.1365....
59.
Gavrylenko, L. M. 2012. New for Ukraine lichens and lichenicolous fungi from the Lower Dnieper. Ukrainian Botanical Journal 69(5): 717–720.
60.
Gavrylenko, L. M. & Khodosovtsev, A. Ye. 2009. Lichens and lichenicolous fungi of the Burgunska balka (Khersonska oblast). Chornomorski Botanical Journal 5: 28–36.
61.
Gizhytska, Z. K. 1929. Material to mycobiota of Ukraine. Bulletin of the Kiev Botanic Garden 10: 4–41.
62.
Gromakova, A. B. 2018. New records of lichens and lichenicolous fungi from Eastern Ukraine. Chornomorski Botanical Journal 14: 269–278. https://doi.org/10.14255/2308-....
63.
Hafellner, J. 1994. Beiträge zu einem Prodromus der lichenicolen Pilze Österreichs und angrenzender Gebiete. I. Einige neue oder seltene Arten. Herzogia 10: 1–28.
64.
Halıcı, M. G. & Candan, M. 2007. Notes on some lichenicolous fungi species from Turkey. Turkish Journal of Botany 31: 353–356.
65.
Halıcı, M. G., Türk, A. Ö. & Candan, M. 2008. Dacampia cladoniicola sp. nov. (Ascomycota, Dacampiaceae) on Cladonia sp. from Turkey. Mycotaxon 103: 53–57. https://doi.org/10.5962/p.4143....
66.
Hawksworth, D. L. 1981. The lichenicolous Coelomycetes. Bulletin of the British Museum for Natural History 9: 1–98.
67.
Hawksworth, D. L. 1992. Nine lichenicolous fungi from Transcarpathians new for Ukraine. Ukrainian Botanical Journal 49(3): 99–101.
68.
Hawksworth, D. L. & Kalb, K. 1992. A new species of Ascochyta on Candelariella aurella from Baja California, Mexico. The Bryologist 95: 338–339. https://doi.org/10.2307/324349....
69.
Hazslinszky, F. 1884. A Magyar birodalom zuzmó-flórája. Kiadja a KM Természettudomanyi Tarsulat.
70.
Hou, L. W., Groenewald, J. Z., Pfenning, L. H., Yarden, O., Crous, P. W. & Cai, L. 2020. The Phoma-like dilemma. Studies in Mycology 96: 309–396. https://doi.org/10.1016/j.simy....
71.
Hou, L. W., Giraldo, A., Groenewald, J. W., Rämä, T., Summerbell, R. C., Huang, G. Z., Cai, L. & Crous, P. W. 2023. Redisposition of acremonium-like fungi in Hypocreales. Studies in Mycology 105: 23–203. https://doi.org/10.3114/sim.20....
72.
Joshi, Y., Bisht, S., Bansal, P. & Pargaien, N. 2024. New additions to lichenicolous mycota of India. Journal of Asia-Pacific Biodiversity 17(2): 290–294. https://doi.org/10.1016/j.japb....
73.
Kapets, N. V. & Kondratyuk, S. Y. 2019. New data on lichenicolous fungi of the Teteriv River Basin (Ukraine). Acta Botanica Hungarica 61: 45–54. https://doi.org/10.1556/034.61....
74.
Katoh, K. & Standley, D. M. 2013. MAFFT Multiple Sequence Alignment Software Version 7: Improvements in Performance and Usability. Molecular Biology and Evolution 30: 772–780. https://doi.org/10.1093/molbev....
75.
Khodosovtsev, A. Ye. 1999. The lichens of the Black Sea steppes. Phytosociocentre, Kyiv.
76.
Khodosovtsev, A. Ye. 2011. A new for Ukraine species of the lichenicolous fungi. Chornomorski Botanical Journal 7(2): 194–198.
77.
Khodosovtsev, A. Ye. 2023. The first records of the lichen-forming, lichenicolous and allied fungi from Ukraine. Folia Cryptogamica Estonica 60: 31–39. https://doi.org/10.12697/fce.2....
78.
Khodosovtsev, A. Ye. & Darmostuk, V. V. 2016a. New species of lichenicolous fungi for Ukraine. Folia Cryptogamica Estonica 53: 93–99. https://doi.org/10.12697/fce.2....
79.
Khodosovtsev, A. Ye. & Darmostuk, V. V. 2016b. Pleospora xanthoriae sp. nov. (Pleosporaceae, Pleosporales), a new lichenicolous fungus on Xanthoria parietina from Ukraine, with a key to the known lichenicolous species of Dacampia and Pleospora. Opuscula Philolichenum 15: 6–11.
80.
Khodosovtsev, A. Ye. & Darmostuk, V. V. 2017a. Zwackhiomyces polischukii sp. nov., and other noteworthy lichenicolous fungi from Ukraine. Polish Botanical Journal 62: 27–35. https://doi.org/10.1515/pbj-20....
81.
Khodosovtsev, A. Ye. & Darmostuk, V. V. 2017b. New for Ukraine records of lichenicolous fungi. Ukrainian Botanical Journal 74: 177–183. https://doi.org/10.15407/ukrbo....
82.
Khodosovtsev, A. Ye. & Darmostuk, V. V. 2018. New for Ukraine species of lichens and lichenicolous fungi from marl limestones in the Northern Black Sea Region. Ukrainian Botanical Journal 75: 33–37. https://doi.org/10.15407/ukrbo....
83.
Khodosovtsev, A. Ye. & Khodosovtseva, Yu. A. 2022. The first contribution to lichens and lichenicolous fungi of Dniprovsko-orilsky Nature Reserve. News Biosphere Reserve “Askania Nova” 24: 36–40. https://doi.org/10.53904/1682-....
84.
Khodosovtsev, A. Ye. & Redchenko, O. O. 2002. An annotated list of the lichen forming fungi of the natural reserve «Mys Martyan» (Ukraine). Ukrainian Botanical Journal 59(1): 64–71.
85.
Khodosovtsev, A. Ye. & Umanets, O. Yu. 2009. Phoma cladoniicola Diederich, Kocourk. & Etayo, a new for Ukraine lichenicolous fungus from Oleshkivsky Sands. Chornomorski Botanical Journal 5(2): 273–275.
86.
Khodosovtsev, A. Ye. & Zavyalova, T. V. 2008. The lichenological zoning of the rock outcrops of the Kayinkulak river shores (Zaporozhskaya oblast, Chernigovsky district). Visnyk of Odessa National University. Biology 13: 56–60.
87.
Khodosovtsev, A. Ye., Vondrák, J. & Šoun, J. 2007. New lichenized and lichenicolous fungi for the Crimean peninsula (Ukraine). Chornomorski Botanical Journal 3: 109–118.
88.
Khodosovtsev, A., Vondrák, J., Naumovich, A., Kocourková, J., Vondráková, O. & Motiejûnaitë, J. 2012. Three new Pronectria species in terricolous and saxicolous microlichen communities (Bionectriaceae, Ascomycota). Nova Hedwigia 95: 211–220. https://doi.org/10.1127/0029-5....
89.
Khodosovtsev, A. Ye., Gavrylenko, L. M. & Klymenko, V. M. 2016a. Katherinomyces cetrariae gen. et sp. nov . (asexual Ascomycota) and Sphaerellothecium aculeatae sp. nov. (Mycosphaerellaceae), new lichenicolous fungi on Cetraria aculeata in Ukraine. Nova Hedwigia 103: 47–55. https://doi.org/10.1127/nova_h....
90.
Khodosovtsev, A. Ye., Darmostuk, V. V. & Nazarchuk, Yu. S. 2016b. Lichens and lichenicolous fungi of the Regional Landscape Park “Tiligulskiy” (Odesa region, Ukraine). Chornomorski Botanical Journal 12: 165–177. https://doi.org/10.14255/2308-....
91.
Khodosovtsev, A., Darmostuk, V., Suija, A. & Ordynets, A. 2018a. Didymocyrtis trassii sp. nov. and other lichenicolous fungi on Cetraria aculeata. The Lichenologist 50: 529–540. https://doi.org/10.1017/S00242....
92.
Khodosovtsev, A. Ye., Darmostuk, V. V., Khodosovtseva, Ya. A., Naumovich, G. O. & Maliuga, N. G. 2018b. The lichens and lichenicolous fungi of the Chalbasy arena in Lower Dnipro sand dunes (Kherson region). Chornomorski Botanical Journal 14: 69–90. https://doi.org/10.14255/2308-....
93.
Khodosovtsev, A. Ye., Darmostuk, V. V., Didukh, Y. P. & Pylypenko, I. O. 2019a. Verrucario viridulae-Staurotheletum hymenogoniae, a new calcicolous lichen community as a component of petrophytic grassland habitats in the Northern Black Sea region. Mediterranean Botany 40: 21–32. https://doi.org/10.5209/MBOT.6....
94.
Khodosovtsev, A. Ye., Darmostuk, V. V., Khodosovtseva, Ya. A. & Gaychenya, Yu. V. 2019b. The lichens and lichenicolous fungi of Trykraty granite massive (Ukraine). Chornomorski Botanical Journal 15: 54–68. https://doi.org/10.32999/ksu19....
95.
Khodosovtsev, O. Ye., Palice, Z., Malíček, J., Svoboda, S., Darmostuk, V., Peksa, O., Bouda, F. & Vondrák, J. 2024. First Ukrainian records of lichens and lichenicolous fungi from Carpathian primeval forests. Czech Mycology 76: 191–213. https://doi.org/10.33585/cmy.7....
96.
Kocakaya, Z., Halıcı, M. G. & Kocakaya, M. 2015. Phoma candelariellae sp. nov., a lichenicolous fungus from Turkey. Mycotaxon 130: 1185–1189. http://dx.doi.org/10.5248/130.....
97.
Kondratyuk, S. Y. 2005. New for mycobiota of Ukraine and rare species of Lichenostigma Haf. (Arthoniales, Ascomycotina). Ukrainian Botanical Journal 62: 509–516.
98.
Kondratyuk, S. Ya., Andrianova, T. V. & Tykhonenko, Yu.Yu. 1999. Study of mycobiota diversity of Ukraine (Lichenicolous, Septoria and Puccinia fungi). Phytosociocentre, Kyiv.
99.
Kondratyuk, S. Y., Khodosovtsev, A. & Kärnefelt, I. 2006. Llimoniella caloplacae sp. nova (Leothiales), a new lichenicolous fungus on Caloplaca borysthenica sp. nova (Lecanorales, Ascomycota). Mycologia Balcanica 3: 95–99.
100.
Kondratyuk, S. Y., Lőkös, L. & Hur, J.-S. 2014. New lichen-forming and lichenicolous fungi from Ukraine. Acta Botanica Hungarica 56: 361–368. https://doi.org/10.1556/ABot.5....
101.
Kondratyuk, S. Y., Upreti, D. K., Mishra, G. K., Nayaka, S., Ingle, K. K., Orlov, O. O., Kondratiuk, A. S., Lőkös, L., Farkas, E., Woo, J.-J. & Hur, J.-S. 2020. New and noteworthy lichen-forming and lichenicolous fungi 10. Acta Botanica Hungarica 62: 69–108. https://doi.org/10.1556/034.62....
102.
Kondratyuk, S. Y., Popova, L. P., Fedorenko, N. M. & Khodosovtsev, O. Y. 2021. Prodromus of sporen plants of Ukraine: Lichens. Naukova dumka, Kyiv.
103.
Krupa, J. 1888. Zapiski mykologiczne z okolic Lwowa i z Podtatrza. Sprawozdanie Komisyji Fizyograficzney Ed. Akademja Umiejętności 22(2): 12–47.
104.
Lawrey, J. D. & Diederich, P. 2003. Lichenicolous fungi: interactions, evolution, and biodiversity. The Bryologist 106: 80–120. https://doi.org/10.1639/0007-2....
105.
Lawrey, J. D., Diederich, P., Nelsen, M. P., Freebury, C., van den Broeck, D., Sikaroodi, M. & Ertz, D. 2012. Phylogenetic placement of lichenicolous Phoma species in the Phaeosphaeriaceae (Pleosporales, Dothideomycetes). Fungal Diversity 55: 195–213. https://doi.org/10.1007/s13225....
106.
Lanfear, R., Calcott, B., Ho, S. Y. W. & Guindon, S. 2012. PartitionFinder: combined selection of partitioning schemes and substitution models for phylogenetic analyses. Molecular Biology and Evolution 29: 1695–1701. https://doi.org/10.1093/molbev....
107.
Lanfear, R., Frandsen, P. B., Wright, A. M., Senfeld, T. & Calcott, B. 2017. PartitionFinder 2: new methods for selecting partitioned models of evolution for molecular and morphological phylogenetic analyses. Molecular Biology and Evolution 34: 772–773. https://doi.org/10.1093/molbev....
108.
Letunic, I. & Bork, P., 2024. Interactive Tree of Life (iTOL) v6: recent updates to the phylogenetic tree display and annotation tool. Nucleic Acids Research 52: W78–W82. https://doi.org/10.1093/nar/gk....
109.
Lypinskyi, V. M., Diachuk, V. A. & Babichenko, V. M. (eds) 2003. Climate of Ukraine: monograph. Raievskyi Publishing House, Kyiv.
110.
Łubek, A., Kukwa, M., Czortek, P. & Jaroszewicz, B. 2019. Lichenicolous fungi are more specialized than their lichen hosts in primeval forest ecosystems, Białowieża Forest, northeast Poland. Fungal Ecology 42: 100866. https://doi.org/10.1016/j.fune....
111.
Malíček, J., Palice, Z., Acton, A., Berger, F., Bouda, F., Sanderson, N. & Vondrák, J. 2018. Uholka primeval forest in the Ukrainian Carpathians – a keynote area for diversity of forest lichens in Europe. Herzogia 31: 140–171. https://doi.org/10.13158/099.0....
112.
Manawasinghe, I. S., Hyde, K. D., Wanasinghe, D. N., Karunarathna, S. C., Maharachchikumbura, S. S. N., Samarakoon, M. C., Voglmayr, H., Pang, K.-L., Chiang, M. W.-L., Jones, E. B. G., Saxena, R. K., Kumar, A., Rajeshkumar, K. C., Selbmann, L., Coleine, C., Hu, Y., Ainsworth, A. M., Liimatainen, K., Niskanen, T., Ralaiveloarisoa, A., Arumugam, E., Kezo, K., Kaliyaperumal, M., Gunaseelan, S., Dissanayake, A. J., Khalid, A. N., Gajanayake, A. J., Flakus, A., Armand, A., Aptroot, A., Rodrigues, A., Tsurykau, A., López-Villalba, Á., De Farias, A. R. G., Sánchez, A., Góes-Neto, A., Goto, B. T., De Souza, C. A. F., Chuaseeharonnachai, C., Lin, C.-G., Li, C., Denchev, C. M., Guerra-Mateo, D., Tennakoon, D. S., Wei, D.-P., Begerow, D., Alves, E., Drechsler-Santos, E. R., Sousa, E. S., De Medeiros, E. V., Langer, E., Zhang, F., De Souza, F. A., Magurno, F., Barreto, G. G., Moreno, G., Mane, G., Alves-Silva, G., Da Silva, G. A., Xia, G., Shen, H.-W., Gui, H., Senanayake, I. C., Luangsa-ard, J. J., Liu, J.-W., Liu, J.-K., Ma, J., Lin, J.-Y., Beserra Jr, J. E. A., Cano-Lira, J. F., Gené, J., Harikrishnan, K., Lu, L., Dos Santos, L. A., Xu, L., Lacerda, L. T., Gusmão, L. F. P., Cáceres, M. E. S., Câmara, M. P. S., De Barros-Barreto, M. B. B., Calabon, M. S., Kukwa, M., Kemler, M., De Melo, M. P., Ghobad-Nejhad, M., Luo, M., Ding, M., Doilom, M., Phonemany, M., Usman, M., Thongklang, N., Boonyuen, N., Ashtekar, N., Kularathnage, N. D., Sruthi, O. P., Kwantong, P., Ansil, P. A., Kooij, P. W., Zhao, Q., Alfenas, R. F., De Oliveira, R. J. V., Singh, R., Da Silva, R. M. F., Avchar, R., Morey, R., Sharma, R., Xu, R.-J., Da Silveira, R. M. B., Xu, R.-F., Jayawardena, R. S., Nanu, S., Nuankaew, S., Tibpromma, S., Boonmie, S., Somrithipol, S., Varghese, S., Moreira, S. I., Rajwar, S., He, S.-C., Kumar, T. K. A., Denchev, T. T., Luangharn, T., De Oliveira, T. G. L., Du, T.-Y., Wen, T.-C., Du, T., Wu, T., Sri-Indrasutdhi, V., Doyle, V. P., Baulin, V., Dong, W., Li, W.-L., Lu, W.-H., Tian, W., Dos Vieira, W. A., Von Brackel, W., Yu, X.-D., Zhang, X., Liu, X.-F., Peng, X.-C., Chen, Y., Yang, Y., Gao, Y., Xiong, Y., Shu, Y., Lu, Y.-Z., Shen, Y.-M., Zhou, Y., Zhang, Y. X., Zhang, W., Luo, Z.-L., Madushani, M. A., Cheewangkoon, R., Song, J. G. & Xu, B. 2025. Fungal diversity notes 1818–1918: taxonomic and phylogenetic contributions on genera and species of fungi. Fungal Diversity 130: 1–261. https://doi.org/10.1007/s13225....
113.
Marynych, O. M. & Shyshchenko, P. H. 2005. Physical geography of Ukraine. Znannia, Kyiv.
114.
Marynych, A. M., Paschenko, M. V. & Shyschenko, P. G. 1985. Landscapes and physico-geographical zoning. In: Marynych, A. M. (ed.), Nature of Ukrainian SSR, pp. 63–72. Naukova dumka, Kyiv.
115.
Minh, B. Q., Schmidt, H. A., Chernomor, O., Schrempf, D., Woodhams, M. D., Von Haeseler, A. & Lanfear, R. 2020. IQ-TREE 2: new models and efficient methods for phylogenetic inference in the genomic era. Molecular Biology and Evolution 37: 1530–1534. https://doi.org/10.1093/molbev....
116.
Nadyeina, O. 2009. The lichen-forming and lichenicolous fungi of the Donetsk Upland (Ukraine). Mycologia Balcanica 6: 37–53.
117.
Nadyeina, O. & Halıcı, M. G. 2012. New lichenicolous fungi records for Kyrgyzstan, Uzbekistan, and Ukraine. Mycotaxon 118: 131–136. https://doi.org/10.5248/118.13....
118.
Naumovich, G. O. & Darmostuk, V. V. 2015. Lichenicolous fungi of the valley of Ingulets river (Ukraine). Chornomorski Botanical Journal 11: 512–520. https://doi.org/10.14255/2308-....
119.
Navarro-Rosinés, P. & Hafellner, J. 1996. Lichenostigma elongata sp. nov. (Dothideales), a lichenicolous ascomycete on Lobothallia and Aspicilia species. Mycotaxon 62: 211–225. https://doi.org/10.5962/p.4159....
120.
Navrotskaya, I. L., Kondratyuk, S. Y., Wasser, S. P., Nevo, E. & Zelenko, S. D. 1996. Lichens and lichenicolous fungi new for Israel and other countries. Israel Journal of Plant Sciences 44: 181–193. https://doi.org/10.1080/079299....
122.
Pem, D., Hyde, K., McKenzie, E., Hongsanan, S., Wanasinghe, D., Boonmee, S., Darmostuk, V., Bhat, J., Tian, Q., Htet, Z., Senanayake, I., Niranjan, M., Sarma, V., Doilom, M. & Dong, W. 2024. A comprehensive overview of genera in Dothideomycetes. Mycosphere 15: 2175–4568. https://doi.org/10.5943/mycosp....
123.
Penn, O., Privman, E., Ashkenazy, H., Landan, G., Graur, D. & Pupko, T. 2010. GUIDANCE: a web server for assessing alignment confidence scores. Nucleic Acids Research 38: W23–W28. https://doi.org/10.1093/nar/gk....
124.
Rehner, S. A. & Samuels, G. J. 1995. Molecular systematics of the Hypocreales: a teleomorph gene phylogeny and the status of their anamorphs. Canadian Journal of Botany 73: 816–823. https://doi.org/10.1139/b95-32....
125.
Ronquist, F., Teslenko, M., van der Mark, P., Ayres, D. L., Darling, A., Höhna, S., Larget, B., Liu, L., Suchard, M. A. & Huelsenbeck, J. P. 2012. MrBayes 3.2: efficient Bayesian phylogenetic inference and model choice across a large model space. Systematic Biology 61: 539–542. https://doi.org/10.1093/sysbio....
126.
Roux, C. & Triebel, D. 1994. Révision des espèces de Stigmidium et de Sphaerellothecium (champignons lichénicoles non lichénisés, Ascomycetes) correspondant à Pharcidia epicymatia sensu Keissler ou à Stigmidium schaereri auct. Bulletin de la Société Linnéenne de Provence 45: 451–542.
127.
Sérusiaux, E., Diederich, P., Brand, A. M. & van den Boom, P. 1999. New or interesting lichens and lichenicolous fungi from Belgium and Luxembourg. VIII. Lejeunia 162: 1–95.
128.
Suija, A., Delhoume, A., Poumarat, S. & Diederich, P. 2021. Didymocyrtis microxanthoriae (Phaeosphaeriaceae, Dothideomycetes), a new lichenicolous fungus from France. Bulletin de la Société des naturalistes luxembourgeois 123: 129–136.
129.
Suija, A., van den Boom, P. P. G., Berger, F., Suija, M., Thayer, J. & Frisch, A. 2026. Taxonomic investigation of Abrothallus (Abrothallales, Ascomycota) species associated with lichen genera Ramalina and Bryoria, including the description of a new species. Nordic Journal of Botany: e05033. https://doi.org/10.1002/njb.05....
130.
Tarieiev, A. S., Girin, A. I., Karpenko, N. I., Tyshchenko, O. V. & Kostikov, I. Yu. 2011. Modified method of DNA extraction from herbarium specimens. Chornomorski Botanical Journal 7(4): 309–317.
131.
Trakunyingcharoen, T., Lombard, L., Groenewald, J. Z., Cheewangkoon, R., Toanun, C., Alfenas, A. C. & Crous, P. W. 2014. Mycoparasitic species of Sphaerellopsis, and allied lichenicolous and other genera. IMA Fungus 5: 391–414. https://doi.org/10.5598/imafun....
132.
Triebel, D. 1989. Lecideicole Ascomyceten. Eine Revision der obligat lichenicolen Ascomyceten auf lecideoiden Flechten. Bibliotheca Lichenologica 35: 1‒278.
133.
Triebel, D. & Scholz, P. 2001. Lichenicolous fungi from Bavaria as represented in the Botanische Staatssammlung München. Sendtnera 7: 211–231.
134.
Triebel, D., Rambold, G. & Elix, J. A. 1995. A conspectus of the genus Phacopsis (Lecanorales). The Bryologist 98: 71–83. https://doi.org/10.2307/324364....
135.
Urbanavichus, G. & Urbanavichene, I. 2014. An inventory of the lichen flora of Lagonaki Highland (NW Caucasus, Russia). Herzogia 27: 285–319. https://doi.org/10.13158/heia.....
136.
van den Boom, P. & Etayo, J. 2014. New records of lichenicolous fungi and lichenicolous lichens from the Iberian Peninsula, with the description of four new species and one new genus. Opuscula Philolichenum 13: 44–79.
137.
van der Kolk, H.-J., Boers, J., Schrier, T., Schrier, N. & Diederich, P. 2024. New lichenicolous fungi from the Netherlands, including the descriptions of Echinothecium micareae and Lichenochora verrucariae. Lindbergia: e026005. https://doi.org/10.25227/linbg....
138.
Varga, N., Lőkös, L. & Farkas, E. 2021. Annotated checklist of the lichenicolous fungi of Hungary. Diversity 13: 557. https://doi.org/10.3390/d13110....
139.
Vondrák, J. & Šoun, J. 2007. Lichenostigma svandae, a new lichenicolous fungus on Acarospora cervina. The Lichenologist 39: 211–216. https://doi.org/10.1017/S00242....
140.
Vouaux, L. 1912. Synopsis des champignons parasites de lichens. Bulletin de la Société Mycologique de France 28: 177–256.
141.
Vouaux, L. 1913. Synopsis des champignons parasites de lichens. Bulletin de la Société Mycologique de France 29: 399–446.
142.
White, T. J., Bruns, T., Lee S. & Taylor, J. 1990. Amplification and direct sequencing of fungal ribosomal RNA genes for phylogenetics. In: Innis, M. A., Gelfand, D. H., Sninsky, J. J. & White, J. W. (eds), PCR Protocols. A Guide to Molecular Methods and Applications. Academic Press, San Diego, pp. 315–322. https://doi.org/10.1016/B978-0....
143.
Zamora, J. C., Millanes, A. M., Wedin, M., Rico, V. J. & Pérez-Ortega, S. 2016. Understanding lichenicolous heterobasidiomycetes: new taxa and reproductive innovations in Tremella s.l. Mycologia 108: 381–396. https://doi.org/10.3852/15-090.
144.
Zelenko, S. D. 2005. Toninia talparum (Lecideaceae, Ascomycetes) – new species for lichenobiota Eurasia from Podoliya Heights. Ukrainian Botanical Journal 62: 517–521.
145.
Zhurbenko, M. P. & Pino-Bodas, R. 2017. A revision of lichenicolous fungi growing on Cladonia, mainly from the Northern Hemisphere, with a worldwide key to the known species. Opuscula Philolichenum 16: 188–266.
146.
Zhurbenko, M., Laursen, G. & Walker, D. A. 2005. New and rare lichenicolous fungi and lichens from the North American Arctic. Mycotaxon 92: 201–212. https://doi.org/10.5962/p.4145....
147.
Zhurbenko, M. P., Enkhtuya, O. & Javkhlan, S. 2019. A first synopsis of lichenicolous fungi of Mongolia, with the description of five new species. Plant and Fungal Systematics 64: 345–366. https://doi.org/10.2478/pfs-20....
148.
Zimmermann, E. & Feusi, S. 2023. Lichenicole Pilze der Schweiz IX: Erwähnenswerte Funde aus dem Mittelland. Meylania 72: 19–34.
149.
Zitenyuk, A. M. & Khodosovtsev, O. Ye. 2024. The genus Solorina (Peltigeraceae, Peltigerales) and its lichenicolous fungi in Ukraine. Chornomorski Botanical Journal 20(4): 378–389. https://doi.org/10.32999/ksu19....
Share
RELATED ARTICLE
| eISSN: | 2657-5000 |
| ISSN: | 2544-7459 |
We process personal data collected when visiting the website. The function of obtaining information about users and their behavior is carried out by voluntarily entered information in forms and saving cookies in end devices. Data, including cookies, are used to provide services, improve the user experience and to analyze the traffic in accordance with the Privacy policy. Data are also collected and processed by Google Analytics tool (more).
You can change cookies settings in your browser. Restricted use of cookies in the browser configuration may affect some functionalities of the website.
You can change cookies settings in your browser. Restricted use of cookies in the browser configuration may affect some functionalities of the website.
